Research focus:
I conduct research on systematics, geography, ecology, and the variability of trees and shrubs in Europe and Mediterranean Sea region.
I am currently dealing with the geographic distribution and conditions for incidence of mountainous trees, shrubs and prostrate shrubs in East Carpathian Mountains in Ukraine.
Also, based on the data in the literature on the subject and my own field observations, I am preparing monographs on selected species of the Juniperus genus which can be found in the Mediterranean Sea region.
In exploring both topics, I collaborate with an extensive group of scientists from Poland and abroad.
Keywords:
trees and shrubs, biodiversity, biogeography, morphology and geographic variability, horology, geographic distribution modelling
Publications
- Boratyński A., Salvà-Catarineu M., Marcysiak K., Mazur M., Romo A., Minissale P., Tan K., Iszkuło G., Witkowski R., Mazur A. 2025. Biology and ecology of the Juniperus phoenicea – J. turbinata – J. canariensis complex II. Genetics, physiology, ecology. Dendrobiology 93, 18–41. https://doi.org/10.12657/denbio.093.002
- Wyka T.P., Robakowski P., Romo A., Boratyński A., Żytkowiak R., Oleksyn J. 2024. Foliar N content parallels increasing aridity in a Mediterranean-Saharan transition zone: Evidence from regional and global trends. Journal of Biogeography 52(1): 213-224. https://doi.org/10.1111/jbi.15027
- Boratyński A., Salvà-Catarineu M., Marcysiak K., Mazur M., Romo Á., Minissale P., Tan K., Iszkuło G., Witkowski R., Mazur A. 2024. Biology and ecology of the Juniperus phoenicea – J. turbinata – J. canariensis complex I. Taxonomy, structure and distribution. Dendrobiology 92: 1-31. https://doi.org/10.12657/denbio.092.001
- Jadwiszczak K.A., Mazur M., Bona A., Marcysiak K., Boratyński A. 2024. Soil requirements, genetic diversity and population history of the Juniperus sabina L. varieties in Europe and Asia. Forests 15(5): 866. https://doi.org/10.3390/f15050866
- Jadwiszczak K., Parzonko S., Mazur M., Marcysiak K., Boratyński A., Bona A. 2024. Ex situ cultivation of the endangered savin junipers (Juniperus sabina L.) from the Western Carpathians. Biodiversity Research and Conservation 75: 27-34. https://doi.org/10.14746/biorc.2024.75.3
- Sękiewicz K., Salva-Catarineu M., Walas Ł., Romo A., Gholizadeh H., Naqinezhad A., Farzaliyev V., Mazur M., Boratyński A. 2024. Consequence of habitat specificity: a rising risk of habitat loss for endemic and sub-endemic woody species under climate change in the Hyrcanian ecoregion. Regional Environmental Change 24(2): 68. https://link.springer.com/article/10.1007/s10113-024-02222-7
- Boratyński A., Dönmez A.A., Dagher-Kharrat M.B., Romo Á., Tan K., Ok T., Iszkuło G., Sobierajska K., Marcysiak K. 2023. Biology and ecology of Juniperus drupacea Labill. Dendrobiology 90: 1-29. https://doi.org/10.12657/denbio.090.001
- Jadwiszczak K.A., Mazur M., Bona A., Marcysiak K., Boratyński A. 2023. Three systems of molecular markers reveal genetic differences between varieties sabina and balkanensis in the Juniperus sabina L. range. Annals of Forest Science 80(1): 45. https://annforsci.biomedcentral.com/articles/10.1186/s13595-023-01211-w
- Lewandowska A., Marcysiak K., Gómez D., Jasińska A.K., Romo A., Didukh Y., Sękiewicz K., Boratyńska K., Boratyński A. 2023. Inference of taxonomic relationships between Rhododendron ferrugineum and R. myrtifolium (Ericaceae) from leaf and fruit morphologies. Botanical Journal of the Linnean Society 201(4): 483-497. https://doi.org/10.1093/botlinnean/boac060
- Tasenkevich L., Boratyński A., Skrypec K., Seniv M., Khmil T., Walas Ł. 2023. Biodiversity of high-mountain woody plants in the East Carpathians in Ukraine. Dendrobiology 89: 1-19. https://doi.org/10.12657/denbio.089.001
- Walas Ł., Pietras M., Mazur M., Romo Á., Tasenkevich L., Didukh Y., Boratyński A. 2023. The perspective of Arctic-Alpine species in southernmost localities: Kalmia procumbens example in the Pyrenees and Carpathians. Plants 12(19): 3399. https://doi.org/10.3390/plants12193399.
- Boratyński A. 2022. Book review: High mountain vascular plants of the Carpathians. Atlas of distribution. Candollea 76(2): 299.https://doi.org/10.15553/c2021v762a11
- Jasińska A.K., Rucińska B., Kozlowski G., Fazan L.,Garfi G., Pasta S., Betrisey S., Gerber E., Safarov H., Van Sam H., Boratyńska K., Manchester S.R., Boratyński A. 2022. Leaf differentiation of extinct and remnant species of Zelkova in Western Eurasia. Plant Biosystems - An International Journal Dealing with all Aspects of Plant Biology 156(6): 1307-1313. https://doi.org/10.1080/11263504.2022.2036849
- Lewandowska A., Boratyńska K., Marcysiak K., Gomez D., Romo A., Malicki M., Iszkuło G., Boratyński A. 2022. Phenotypic differentiation of Rhododendron ferrugineum populations in European mountains. Dendrobiology 87: 1-12. https://doi.org/10.12657/denbio.087.001
- Boratyńska K., Gołąb Z., Łabiszak B., Niemczyk W., Sobierajska K.I., Ufnalski K., Wachowiak W, Boratyński A. 2021. Are there any traces of Pinus uliginosa in the Stołowe Mountains outside the Wielkie Tofowisko Batorowskie and Błędne Skały? Acta Societatis Botanicorum Poloniae. 90:904. https://doi.org/10.5586/asbp.904
- Boratyńska K., Jasińska A.K., Sobierajska K.I., Salva Catarineu M., Romo A., Boratyński A. 2021. Morphology supports the geographic pattern of genetic differentiation of Pinus sylvestris (Pinaceae) in the Iberian Peninsula. Plant Biosystems. https://doi.org/10.1080/11263504.2021.1922532
- Jasińska A.K., Rucińska B., Kozlowski G., Fazan L., Garfì G., Pasta S., Bétrisey S., Gerber E., Boratyńska K., Boratyński A. 2021. Taxonomic relationships and population differentiation of the south-western Eurasian Zelkova species inferred in leaf morphology. Dendrobiology 85: 60–77, https://doi.org/10.12657/denbio.085.007
- Litkowiec M., Sękiewicz K., Romo A., Ok T., Bou Dagher-Kharrat M., Jasińska A.K., Sobierajska K., Boratyńska K., Boratyński A. 2021. Biogeography and relationships of the Abies taxa from the mediterranean and central Europe regions as revealed by nuclear DNA markers and needle structural characters. Forest Ecology and Management 479: #118606. https://doi.org/10.1016/j.foreco.2020.118606
- Mazur M., Boratyński A., Boratyńska K., Marcysiak K. 2021. Weak geographical structure of Juniperus sabina (Cupressaceae) morphology despite its discontinuous range and genetic differentiation. Diversity 13: 470. https://doi.org/10.3390/d13100470
- Salvà-Catarineu M., Romo A., Mazur M., Zielińska M., Minissale P., Dönmez A.A., Boratyńska K., Boratyński A. 2021. Past, present and future geographic range of the relict Mediterranean and Macaronesian Juniperus phoenicea complex. Ecology and Evolution 11: 5075–5095, https://doi.org/10.1002/ece3.7395
- Song Y-G., Walas Ł., Pietras M., Sâm H.V., Yousefzadeh H., Ok T., Farzaliyev V., Worobiec G., Worobiec E., Stachowicz-Rybka R., Kozlowski G., Boratyński A., Boratyńska K., Jasińska A.K. 2021. Past, present and future suitable areas for the relict tree Pterocarya fraxinifolia (Juglandaceae): integrating fossil records, niche modelling, and phylogeography for conservation. European Journal of Forest Research. https://doi.org/10.1007/s10342-021-01397-6
- Sobierajska K., Wachowiak W., Zaborowska J., Łabiszak B., Wójkiewicz B., Sękiewicz M., Jasińska A.K., Sękiewicz K., Boratyńska K., Marcysiak K., Boratyński A. 2020.Genetic consequences of hybridization in relict isolated trees Pinus sylvestris and the Pinus mugo complex. Forests 11: #1086. https://doi.org/10.3390/f11101086
- Taib A., Morsli A., Chojnacka A., Walas Ł., Sękiewicz K., Boratyński A., Romo À., Dering M. 2020. Patterns of genetic diversity in North Africa: Moroccan-Algerian genetic split in Juniperus thurifera subsp. africana. Scientific Reports 10: 1-17. https://doi.org/10.1038/s41598-020-61525-x
- Boratyńska K., Tomaszewski D., Montserrat J.M., Marek S., Boratyński A. 2019. Taxonomic position of Pinus ceciliae (Pinaceae) endemic for Balearic Islands as revealed on needle characteristics. Dendrobiology 82: 8-16. http://dx.doi.org/10.12657/denbio.082.002
- Kosiński P., Sękiewicz K., Walas Ł., Boratyński A., Dering M. 2019. Spatial genetic structure of the endemic alpine plant Salix serpillifolia: genetic swamping on nunataks due to secondary colonization? Alpine Botany 129: 107-121. https://doi.org/10.1007/s00035-019-00224-4
- Kosiński P., Sliwinska E., Hilpold A., Boratyński A. 2019. DNA ploidy in Salix retusa agg. only partly in line with its morphology and taxonomy. Nordic Journal of Botany 37: #e02197. https://doi.org/10.1111/njb.02197
- Romo A., Mazur M., Salvà-Catarineu M., Boratyński A. 2019. A re-evaluated taxon: Genetic values and morphological characters support the recognition of the Canary Island juniper of the phoenicea group at a specific level. Phytotaxa 406: 64-70. https://doi.org/10.11646/phytotaxa.406.1.3
- Walas Ł., Sobierajska K., Ok T., Dönmez A.A., Kanoğlu S.S., Dagher-Kharrat M.B., Douaihy B., Romo A., Stephan J., Jasińska A.K., Boratyński A. 2019. Past, present, and future geographic range of an oro-Mediterranean Tertiary relict: The Juniperus drupacea case study. Regional Environmental Change 19: 1507-1520. https://doi.org/10.1007/s10113-019-01489-5
- Adams R.P., Boratyński A., Marcysiak K., Roma-Marzio F., Peruzzi L., Bartolucci F., Conti F., Mataraci T., Schwarzbach A.E., Tashev A.N., Siljak Yakovlev S. 2018. Discovery of Juniperus sabina balkanensis R. P. Adams and A. N. Tashev in Macedonia, Bosnia-Herzegovina, Croatia and Central and Southern Italy and relictual polymorphism found in nrDNA. Phytologia 100: 117-127.
- Mazur M., Zielińska M., Boratyńska K., Romo A., Salva-Catarineu M., Marcysiak K., Boratyński A. 2018. Taxonomic and geographic differentiation of Juniperus phoenicea agg. based on cone, seed, and needle characteristics. Systematics and Biodiversity 16: 469-482. https://doi.org/10.1080/14772000.2018.1439120
- Sękiewicz K., Dering M., Romo A., Bou Dagher-Kharrat M., Boratyńska K., Ok T., Boratyński A. 2018. Phylogenetic and biogeographic insights of long-lived Mediterranean Cupressus taxa with a schizo-endemic distribution and Tertiary origin. Botanical Journal of the Linnean Society 188: 190-212. https://doi.org/10.1093/botlinnean/boy049
- Adams R.P., Boratyński A., Dagher-Kharrat M.B., Leschner H., Mataraci T. 2017. Geographic variation in Juniperus drupacea: DNA sequencing and volatile leaf oils: Further evidence of putative Pleistocene genetic isolation between Europe and Asia. Phytologia 99: 249-257.
- Adams R. P., Boratyński A., Mataraci T., Tashev A. N., Schwarzbach A. E. 2017. Discovery of Juniperus sabina balkanensis R. P. Adams and A. N. Tashev in western Turkey. Phytologia 99: 22-31.
- Dering M., Kosiński P., Wyka T.P., Pers-Kamczyc E., Boratyński A., Boratyńska K., Reich P.B., Romo A., Zadworny M., Żytkowiak R., Oleksyn J. 2017. Tertiary remnants and Holocene colonizers: Genetic structure and phylogeography of Scots pine reveal higher genetic diversity in young boreal than in relict Mediterranean populations and a dual colonization of Fennoscandia. Diversity and Distributions 23: 540-555. https://doi.org/10.1111/ddi.12546
- Dzialuk A., Boratyńska K., Romo A., Boratyński A. 2016. Taxonomic and geographic variation of the Pinus mugo complex on chloroplast microsatellite markers. Systematics and Biodiversity 15: 464-479. https://doi.org/10.1080/14772000.2016.1257518
- Dering M., Latałowa M., Boratyńska K., Kosiński P., Boratyński A. 2017. Could clonality contribute to the northern survival of grey alder [Alnus incana (L.) Moench] during the Last Glacial Maximum? Acta Societatis Botanicorum Poloniae 86: #3523. https://doi.org/10.5586/asbp.3523
- Jasińska A., Boratyńska K., Sękiewicz K., Gristina E.D., Boratyński A. 2017. Relationships among Abies nebrodensis, A. alba and A. cephalonica in the morphological and anatomical needle characteristics. Plant Biosystems - An International Journal Dealing with all Aspects of Plant Biology 151: 775-782. https://doi.org/10.1080/11263504.2017.1311960
- Jasińska A.K., Sękiewicz K., Ok T., Romo A., Boratyński A., Boratyńska K. 2017. Taxonomic position of Abies equi-trojani on the basis of needle characters by comparing with different fir species. Turkish Journal of Botany 41: 620-631. https:// https://journals.tubitak.gov.tr/botany/vol41/iss6/9/
- Kosiński P., Boratyński A., Hilpold A. 2017. Taxonomic differentiation of Salix retusa (Salicaceae) based on leaf characteristics. Dendrobiology 78: 40-50. http://dx.doi.org/10.12657/denbio.078.005
- Romo A., Iszkuło G., Seghir Taleb M., Walas Ł., Boratyński A. 2017. Taxus baccata in Morocco: a tree in regression in its southern extreme. Dendrobiology 78: 63-74. http://dx.doi.org/10.12657/denbio.078.007
- Adams R.P., Armagan M., Boratyński A., Douaihy B., Dagher-Kharrat M.D., Farzaliyev V., Gucel S., Mataraci T., Tashev A.N., Schwarzbach A.E. 2016. Evidence of relictual introgression or incomplete lineage sorting in nrDNA of Juniperus excelsa and polycarpos in Asia Minor. Phytologia 98(2): 146-155.
- Mazur M., Minissale P., Sciandrello S., Boratyński A. 2016. Morphological and ecological comparison of populations of Juniperus turbinata Guss. and J. phoenicea L. from the Mediterranean region. Plant Biosystems 150: 313-322. https://doi.org/10.1080/11263504.2014.994579
- Sękiewicz K., Boratyńska K., Dagher-kharrat M.B., Ok T., Boratyński A. 2016. Taxonomic differentiation of Cupressus sempervirens and C. atlantica based on morphometric evidence. Systematics and Biodiversity 15(5): 494-508. https://doi.org/10.1080/14772000.2016.1171260
- Sobierajska K., Boratyńska K., Jasińska A., Dering M., Ok T., Douaihy B., Bou Dagher-Kharrat M., Romo Á., Boratyński A. 2016. Effect of the Aegean Sea barrier between Europe and Asia on differentiation in Juniperus drupacea (Cupressaceae). Botanical Journal of the Linnean Society 180: 365-385. https://doi.org/10.1111/boj.12377
- Boratyńska K., Jasińska A.K., Boratyński A. 2015. Taxonomic and geographic differentiation of Pinus mugo complex on the needle characteristics. Systematics and Biodiversity 13(6): 581-595. https://doi.org/10.1080/14772000.2015.1058300
- Boratyńska K., Sękiewicz K., Jasińska A.K., Tomaszewski D., Iszkuło G., Ok T., Bou Dagher-Kharrat M., Boratyński A. 2015. Effect of geographic range discontinuity on taxonomic differentiation of Abies cilicica. Acta Societatis Botanicorum Poloniae 84(4): 419-430. https://doi.org/10.5586/asbp.2015.037
- Jasińska A.K., Rucińska B., Kozlowski G., Bétrisey S., Safarov H., Boratyńska K., Boratyński A. 2015. Morphological differentiation of leaves in the relict tree Zelkova carpinifolia (Ulmaceae). Dendrobiology 74: 109-122. http://dx.doi.org/10.12657/denbio.074.011
- Sękiewicz K., Dering M., Sękiewicz M., Boratyńska K., Iszkuło G., Litkowiec M., Ok T., Dagher-Kharrat M.B., Boratyński A. 2015. Effect of geographic range discontinuity on species differentiation—East-Mediterranean Abies cilicica: a case study. Tree Genetics & Genomes 11: #810. https://doi.org/10.1007/s11295-014-0810-5
- Adams R.P., Arista M., Boratyński A., Houari H.H., Leschner H., Liber Z., Minissale P., Sciandrello S., Mataraci T. 2014. Geographic variation in the leaf essential oil of Juniperus turbinata from throughout its range in the Mediterranean. Phytologia 96: 149-158.
- Boratyńska K., Dzialuk A., Lewandowski A., Marcysiak K., Jasińska A.K., Sobierajska K., Tomaszewski D., Burczyk J., Boratyński A. 2014. Geographic distribution of quantitative traits variation and genetic variability in natural populations of Pinus mugo in Central Europe. Dendrobiology 72: 65-84. http://dx.doi.org/10.12657/denbio.072.006
- Boratyński A., Wachowiak W., Dering M., Boratyńska K., Sękiewicz K., Sobierajska K., Jasińska A.K., Klimko M., Montserrat J.M., Romo A., Ok T., Didukh Y. 2014. The biogeography and genetic relationships of Juniperus oxycedrus and related taxa from the Mediterranean and Macaronesian regions. Botanical Journal of the Linnean Society 174(4): 637-653. https://doi.org/10.1111/boj.12147
- Dering M., Sękiewicz K., Boratyńska K., Litkowiec M., Iszkuło G., Romo A., Boratyński A. 2014. Genetic diversity and inter-specific relations of western Mediterranean relic Abies taxa as compared to the Iberian A. alba. Flora 209(7): 367-374 https://doi.org/10.1016/j.flora.2014.03.011
- Jasińska A.K., Boratyńska K., Dering M., Sobierajska K.I., Ok. T., Romo A., Boratyński A. 2014. Distance between south-European and south-west Asiatic refugial areas involved morphological differentiation – Pinus sylvestris case study. Plant Systematics and Evolution 300: 1487-1502. https://doi.org/10.1007/s00606-013-0976-6
- Romo A., Boratyński A. 2014. The genus Cotoneaster (Rosaceae) in NW Africa. Willdenowia 44: 229-239. https://doi.org/10.3372/wi.44.44204
- Sękiewicz K., Sękiewicz M., Romo A., Didukh Y., Fennane M., Boratyński A. 2014. Chorological and conservation status of the endemic cypress, Cupressus atlantica Gaussen, in the High Atlas (Morocco). Dendrobiology 71: 3-13. http://dx.doi.org/10.12657/denbio.071.001
- Boratyński A., Jasińska A.K., Marcysiak K., Mazur M., Romo A.M., Boratyńska K., Sobierajska K., Iszkuło G. 2013. Morphological differentiation supports the genetic pattern of the geographic structure of Juniperus thurifera (Cupressaceae). Plant Systematics and Evolution 299: 773-784. https://doi.org/10.1007/s00606-013-0760-7
- Jasińska A.K., Boratyńska K., Sobierajska K., Romo A., Ok. T., Bou Dagher-Kharat M., Boratyński A. 2013. Relationships among Cedrus libani, C. brevifolia and C. atlantica as revealed by the morphological and anatomical needle characters. Plant Systematics and Evolution 299: 35-48. https://doi.org/10.1007/s00606-012-0700-y
- Romo A., Hidalgo O., Boratyński A., Sobierajska K., Jasińska A.K., Valles J., Garnatje T. 2013. Genome size and ploidy levels in highly fragmented habitats: the case of western Mediterranean Juniperus (Cupressaceae) with special emphasis on J. thurifera L. Tree Genetics & Genomes 9: 587-599. https://doi.org/10.1007/s11295-012-0581-9
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• Sękiewicz K., Sękiewicz M., Jasińska A.K., Boratyńska K., Iszkuło G., Romo A., Boratyński A. 2013. Morphological diversity and structure of West Mediterranean Abies species. Plant Biosystems 147(1): 125-134. https://doi.org/10.1080/11263504.2012.753130
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Douaihy B., Sobierajska K., Jasińska A.K., Boratyńska K., Ok T., Romo A., Machon N., Didukh Y., Bou Dagher-Kharrat M., Boratyński A. 2012. Morphological versus molecular markers to describe variability in Juniperus excelsa subsp. excelsa (Cupressaceae). Annals of Botany Plants #pls013. https://doi.org/10.1093/aobpla/pls013
- Dzialuk A., Boratyński A., Boratyńska K., Burczyk J. 2012. Geographic patterns of genetic diversity of Pinus mugo (Pinaceae) in Central European mountains. Dendrobiology 68: 31-41. https://www.idpan.poznan.pl/pl/vol-68/68-31-41
- Iszkuło G., Golimowski R., Lewandowska A., Wachowiak E., Boratyński A. 2012. Changes in vegetation of the ‘Cisy Staropolskie im. Leona Wyczółkowskiego’ reserve near Wierzchlas in the Tuchola Forest. Sylwan 156: 163-169.
- Boratyński A., Dubicki A., Iszkuło G., Jasińska A.K., Boratyńska K., Piórkowska M., Marcysiak K. 2011. How environmental signals affect frequency of three-needle dwarf shoots on Pinus mugo. Polish Journal of Environmental Studies 20(1): 45-52.
- Douaihy B., Vendramin G.G., Boratyński A., Machon N., Bou Dagher-Kharrat M. 2011. High genetic diversity with moderate differentiation in Juniperus excelsa from Lebanon and the East Mediterranean region. Annals of Botany, Plants Advance Access 21: 1-34 https://doi.org/10.1093/aobpla/plr003
- Dzialuk A., Mazur M., Boratyńska K., Montserrat J.M., Romo A., Boratyński A. 2011. Population genetic structure of Juniperus phoenicea (Cupressaceae) in the western Mediterranean Basin: gradient of diversity on a broad geographical scale. Annals of Forest Science 68: 1341-1350. https://doi.org/10.1007/s13595-011-0150-7
- Iszkuło G., Boratyński A. 2011. Initial period of sexual maturity determines the greater growth rate of male over female in the dioecious tree Juniperus communis subsp. communis. Acta Oecologica-International Journal of Ecology 37: 99-102. https://doi.org/10.1016/j.actao.2011.01.001
- Boratyński A., Marcysiak K., Lewandowska A., Jasińska A., Iszkuło G. 2010. Interrelations among con-generic and co-occurring tree species: asymmetric hybridization and the high success of Quercus petraea (Matt.) Liebl. regeneration in mixed Q.petraea/Q.robur L. stands. Polish Journal of Ecology 58(2): 273-283.
- Jasińska A.K., Wachowiak W., Muchewicz E., Boratyńska K., Montserrat J.M., Boratyński A. 2010. Cryptic hybrids between Pinus uncinata and P. sylvestris. Botanical Journal of the Linnean Society 163: 473-485. https://doi.org/10.1111/j.1095-8339.2010.01065.x
- Mazur M., Klajbor K., Kielich M., Sowińska M., Romo A., Montserrat J.M., Boratyński A. 2010. Intra-specific differentiation of Juniperus phoenicea in the western Mediterranean region revealed in morphological multivariate analysis. Dendrobiology 63: 21-31. https://www.idpan.poznan.pl/pl/vol-63/63-21-31
- Romo A.M., Boratyński A. 2010. A New combination in Erica (Ericaceae). Collectanea Botanica 29: 95-98.
- Boratyński A., Jasińska A., Boratyńska K., Iszkuło G., Piórkowska M. 2009. Life span of needles of Pinus mugo Turra: effect of altitude and species origin. Polish Journal of Ecology 57(3): 567-572.
- Boratyński A., Lewandowski A., Boratyńska K., Montserrat J.M., Romo A. 2009. High level of genetic differentiation of Juniperus phoenicea (Cupressaceae) in the Mediterranean region: geographic implications. Plant Systematics and Evolution 277: 163-172. https://doi.org/10.1007/s00606-008-0122-z
- Dzialuk A., Muchewicz E., Boratyński A., Montserrat J.M., Boratyńska K., Burczyk J. 2009. Genetic variation of Pinus uncinata (Pinaceae) in the Pyrenees determined with cpSSS markers. Plant Systematics and Evolution 277: 197-205. https://doi.org/10.1007/s00606-008-0123-y
- Iszkuło G., Jasińska A.K., Giertych M. J., Boratyński A. 2009. Do secondary sexual dimorphism and female intolerance to drought influence the sex ratio and extintion risk of Taxus baccata? Plant Ecology 200: 229-240. https://doi.org/10.1007/s11258-008-9447-5
- Boratyński A., Marcysiak K., Lewandowska A., Jasińska A., Iszkuło G., Burczyk J. 2008. Differences in leaf morphology between Quercus petraea and Q. robur adult and young individuals. Silva Fennica 42(1): 115-124. https://doi.org/10.14214/sf.268
- Boratyńska K., Boratyński A. 2007. Taxonomic differences among closely related pines Pinus sylvestris, P. mugo, P. uncinata, P. rotundata and P. uliginosa as revealed in needle sclerenchyma cells. Flora 202: 555-569. https://doi.org/10.1016/j.flora.2006.11.004
- Boratyński A., Boratyńska K., Mazur M., Marcysiak K. 2007. Seed involucre variation in Carpinus betulus (Corylaceae) in Poland. Acta Biologica Cracoviensia, series Botanica. 49(1): 103-111.
- Klimko M., Boratyńska K., Montserrat J.M., Didukh Ya., Romo A., Gómez D., Kluza-Wieloch M., Marcysiak K., Boratyński A. 2007. Morphological variation of Juniperus oxycedrus ssp. oxycedrus (Cupressaceae) in the Mediterranean region. Flora 202: 133-147. https://doi.org/10.1016/j.flora.2006.03.006
- Marcysiak K., Boratyński A. 2007. Contribution to the taxonomy of Pinus uncinata (Pinaceae) based on cone characters. Plant Systematics and Evolution 264: 57-73. https://doi.org/10.1007/s00606-006-0501-2
- Marcysiak K., Mazur M., Romo A., Montserrat J.M., Didukh Y., Boratyńka K., Jasińska J., Kosiński P., Boratyński A. 2007. Numerical taxonomy of Juniperus thurifera, J. excelsa and J. foetidissima (Cupressaceae) based on morphological characters. Botanical Journal of the Linnean Society 155: 483-495. https://doi.org/10.1111/j.1095-8339.2007.00730.x
- Boratyński A., Piwczyński M., Didukh Y.P., Tasenkevich L., Romo A., Ratyńska H. 2006. Distribution and phytocoenotic characteristics of relict populations of Rhododendron myrtifolium (Ericaceae) in the Ukrainian Carpathians. Polish Botanical Studies 22: 53-62.
- Tsaryk I., Didukh Ya. P., Tasenkevich L., Waldon B., Boratyński A. 2006. Pinus mugo Turra (Pinaceae) in the Ukrainian Carpathians. Dendrobiology 55: 39–49.
- Ratyńska H., Waldon B., Boratyński A. 2006. Xerothermic and anthropophytic species populations in the grassland communities of 'Skarpy Ślesińskie" Nature Reserve and its surroundings. Biodiversity Research and Conservation. 3-4: 381-385.
- Romo A., Boratyński A. 2007. Nomenclatural note on Juniperus thurifera subsp. africana (Cupressaceae). Annales Botanici Fennici 44(1): 72-75.
- Romo A., Boratyński A. 2007. Notes and contributions to the vascular flora of Oukaimeden (Central High Atlas, Morocco). Candollea 62(1): 69-90.
- Waldon B., Ratyńska H., Boratyński A. 2006. Differentiation of the plant cover on slopes of the Noteć valley near Ślesin (northern Poland). Biodiversity Research and Conservation 3-4: 348-351.
- Boratyńska K., Boratyński A. 2006. Occurrence of three-needle dwarf shoots on European species of genus Pinus (Pinaceae). Plant Biosystems 140(1): 21-26. https://doi.org/10.1080/11263500500500796
- Iszkulo G., Boratynski A. 2006. Analysis of the relationship between photosynthetic photon flux density and natural Taxus baccata seedlings occurrence. Acta Oecologica 29(1): 78-84. https://doi.org/10.1016/j.actao.2005.08.001
- Lewandowski A., Burczyk J., Wachowiak W., Boratyński A., Prus-Głowacki W. 2005. Genetic evaluation of seeds of highly endangered Pinus uliginosa Neumann from Węgliniec reserve for ex-situ conservation program. Acta Societatis Botanicorum Poloniae 74(3): 237-242.
- Romo A., Boratyński A. 2005. Chorology of Juniperus thurifera (Cupressaceae) in Morocco. Dendrobiology 54: 41-50.
- Boratyńska K., Marcysiak K., Boratyński A. 2005. Pinus mugo (Pinaceae) in the Abruzzi Mountains: high morphological variation in isolated populations. Botanical Journal of the Linnean Society 147: 309-319. https://doi.org/10.1111/j.1095-8339.2005.00374.x
- Iszkuło G., Boratyński A. 2005. Different age and spatial structure of two spontaneous subpopulations of Taxus baccata as a result of various intensity of colonization process. Flora 200 (2): 195-206. https://doi.org/10.1016/j.flora.2004.03.001
- Iszkuło G., Boratyński A., Didukh Y., Romaschenko K., Pryazhko N. 2005. Changes of population structure of Taxus baccata during 25 years in protected area (the Carpathian Mountains, East Ukraine). Polish Journal of Ecology 53: 13-23.
- Didukh Ya., Romo A., Boratyński A. 2004. On five rare vascular plant species reported from Crimea, Ukraine. Willdenowia 34: 1-4.
- Iszkuło G., Boratyński A. 2004. Interaction between canopy tree species and European yew Taxus baccata (Taxaceae). Polish Journal of Ecology 52(4): 501-509.
- Klimko M., Boratyńska K., Boratyński A., Marcysiak K. 2004. Morphological variation of Juniperus oxycedrus subsp. macrocarpa (Cupressaceae) in three Italian localities. Acta Societatis Botanicorum Poloniae 73: 113-119. https://doi.org/10.5586/asbp.2004.016
- Mazur M., Boratyńska K., Marcysiak K., Didukh Y., Romo A., Kosiński P., Boratyński A. 2004. Low level of inter-populational differentiation in Juniperus excelsa M. Bieb. (Cupressaceae). Dendrobiology 52: 39-46. https://www.idpan.poznan.pl/pl/vol-52/52-39-46
- Romo A., Didukh Y., Boratyński A. 2004. Thesium (Santalaceae) in Crimea, Ukraine. Annales Botanici Fennici 41: 273-281.
- Boratyńska K., Boratyński A. 2003. Anomalous needle numbers on dwarf shoots of Pinus mugo and P. uncinata (Pinaceae). Acta Societatis Botanicorum Poloniae 72: 139-148. https://doi.org/10.5586/asbp.2003.019
- Boratyńska K., Boratyński A., Lewandowski A. 2003. Morphology of Pinus uliginosa (Pinaceae) needles from populations exposed to and isolated from the direct influence of Pinus sylvestris. Botanical Journal of the Linnean Society 142: 83-91. https://doi.org/10.1046/j.1095-8339.2003.00156.x
- Boratyński A., Boratyńska K., Lewandowski A., Gołąb Z., Kiciński P. 2003. Evidence of the possibility of natural reciprocal crosses between Pinus sylvestris and P. uliginosa based on the phenology of reproductive organs. Flora 198: 377-388. https://doi.org/10.1078/0367-2530-00110
- Boratyński A., Lewandowska A., Ratyńska H. 2003. Cerasus fruticosa Pall. (Rosaceae) in the region of Kujavia and South Pomerania (N Poland). Dendrobiology 49: 3-13. https://www.idpan.poznan.pl/pl/vol-499/49-03-13
- Boratyński A., Ratyńska H., Waldon B., Bartczak A. 2003. Flora and vegetation of walls in the town of Krosno Odrzańskie (Poland). Collectanea Botanica (Barcelona) 26: 129-139.
- Boratyński A., Romo A. 2003. Loiseleuria procumbens (Ericaceae) in the Spanish Pyrenees. Acta Societatis Botanicorum Poloniae 72: 125-133. https://doi.org/10.5586/asbp.2003.017
- Mazur M., Boratyńska K., Marcysiak K., Gómez D., Tomaszewski D., Didukh J., Boratyński A. 2003. Morphological variability of Juniperus phoenicea (Cupressaceae) from three distant localities on Iberian Peninsula. Acta Societatis Botanicorum Poloniae 72(1): 71-78. https://doi.org/10.5586/asbp.2003.009
- Boratyński A., Didukh Ya.P. 2002. Loiseleuria procumbens (Ericaceae) in the Ukrainian Carpathians. Dendrobiology 47: 3-8. https://www.idpan.poznan.pl/pl/vol-477/47-03-08
- Didukh Ya. P., Boratyński A. 2002. The genus Celtis L. (Ulmaceae) in flora of Ukraine. Ukrayins'kyi Botanichnyi Zhurnal 59(1): 5-9.
- Lewandowski A., Samoćko J., Boratyńska K., Boratyński A. 2002. Genetic diferrences between two Polish populations of Pinus uliginosa, compared to P. sylvestris and P. mugo. Dendrobiology 48: 51-57. https://www.idpan.poznan.pl/pl/vol-488/48-51-57
- Świerkosz K., Boratyński A. 2002. Chorological and synanthropodynamical analysis of trees and shrubs of the Stołowe Mts. (Middle Sudety). Dendrobiology 48: 75-85. https://www.idpan.poznan.pl/pl/vol-488/48-75-85
- Boratyński A., Didukh Ya., Lucak M. 2001. The yew (Taxus baccata L.) population in Knyazhdvir Nature Reserve in the Carpathians (Ukraine). Dendrobiology 46: 3-8. https://www.idpan.poznan.pl/pl/vol-466/46-03-08
- Lewandowski A., Boratyński A., Mejnartowicz L. 2000. Allozyme investigations on the genetic differentiation between closely related pines – Pinus sylvestris, P. mugo, P. uncinata and P. uliginosa (Pinaceae). Plant Systematic and Evolution 221: 15-24.
- Lewandowski A., Samoćko J., Boratyński A., Mejnartowicz L. 2000. Inheritance and linkage of allozymes in Juniperus phoenicea L. (Cupressaceae). Acta Societatis Botanicorum Poloniae 69 (3): 201-205.
- Boratyński A., Didukh Ya. P. 2000. The genus Empetrum (Empetraceae) in Ukraine. Fragmenta Floristica et Geobotanica 45(1–2): 423-431.
Publications before year 2000 you can find here: https://www.scopus.com/authid/detail.uri?authorId=6603028980
Projects
- Subalpine and Alpine woody plants in the Eastern Carpathians of Ukraine. (Original title: Zdrewniałe rośliny subalpejskie i alpejskie w Karpatach Wschodnich Ukrainy). Project funded by the Internal Research Fund of the Institute of Dendrology, Polish Academy of Sciences (FBW_4, DEC-2024/04/ZB/FBW/02). Project duration: 2024–2025. Principal Investigator: Boratyński A.
- Assessment of the contribution of Peat-bog Pine (Pinus uliginosa) genes in Scots Pine (Pinus sylvestris) populations in the Stołowe Mountains National Park. (Original title: Ocena udziału genów sosny błotnej (Pinus uliginosa) w populacjach sosny zwyczajnej (Pinus sylvestris) na terenie Parku Narodowego Gór Stołowych). Project funded by the Forest Fund of the State Forests (ZBN/03/2020). Project duration: Principal Investigator: Boratyński A.
- Geographical differentiation of Juniperus oxycedrus and related taxa from the Mediterranean refugia of Pleistocene flora. (Original title: Zróżnicowanie geograficzne Juniperus oxycedrus L. i taksonów pokrewnych z obszaru śródziemnomorskich ostoi flor plejstoceńskich). Project funded by the Ministry of Science and Higher Education (N N303 1534 33). Project duration: 2007–2010. Principal Investigator: Boratyński A.
- Differentiation of Pinus uncinata Ramond (Pinaceae) within the natural range. (Original title: Zróżnicowanie Pinus uncinata Ramond (Pinaceae) w granicach zasięgu naturalnego). Project funded by the Scientific Research Committee – Ministry of Science and Higher Education (2P04C 018 29). Project duration: 2005–2007. Principal Investigator: Boratyński A.
- Silver fir (Abies alba Mill) in the Sudetes - characteristics of fragments of endangered populations. (Original title: Jodła pospolita (Abies alba Mill) w Sudetach - charakterystyka fragmentów zagrożonych populacji). Project funded by the Regional Directorate of State Forests in Wrocław. Project duration: 1998–2001. Principal Investigator: Boratyński A.
Popular Science Articles
[Articles in Polish]
- Boratyński A., Boratyńska K. 2024. Jałowiec pospolity (Juniperus communis) – uwagi dotyczące przyszłości gatunku w Polsce. Rocznik Polskiego Towarzystwa Dendrologicznego 70 (2022/2023): 27-35.
- Boratyński A., Boratyńska K. 2022. Jałowce występujące w sposób naturalny w Europie i w regionie śródziemnomorskim – przegląd. Rocznik Polskiego Towarzystwa Dendrologicznego 69: 17-34.
Books and Book Chapters
- Petrova A., Boratyński A. 2022. Book reviews. Phytologia Balcanica 27(3): 413 – 419.
- Adams R.P., Boratyński A., Marcysiak K., Roma-Marzio F., Peruzzi L., Bartolucci F., Conti F., Mataraci T., Schwarzbach A.E., Tashev A.N., Siljak Yakovlev S. 2018. Discovery of Juniperus sabina balkanensis R. P. Adams and A. N. Tashev in Macedonia, Bosnia-Herzegovina, Croatia and Central and Southern Italy and relictual polymorphism found in nrDNA. Phytologia 100: 117-127.
- Didukh Ya., Kontar I., Boratyński A. 2018. Phytoindicating comparison of vegetation of the Polish Tatras, the Ucrainian Carpathians, and the Mountain Crimea. In: Greller A.M., Fujiwara K., Pedrotti F. (eds), Geographical changes in vegetation and plant functional types, geobotany studies. Springer. pp. 186-210.
- Boratyński A. 2007. The Central-European disjunction in the range of Norway spruce. In: Tjoelker M., Boratyński A., Bugała W. (eds.). Biology and ecology of Norway spruce. Springer, Forestry Sciences 78: 37-47.
- Boratyński A. 2002. Loranthaceae (Viscaceae). In: Valdes B., Rejdali M., Achhal A. El Kadmiri, Jury J.L., Montserrat J.M. (eds.). Checklist of Vascular Plants of Morocco with identification keys. Consejo Superior de Investigaciones Cientificas. Madrid. 1: 410-411.
- Boratyński A. 2002. Oleaceae. In: Valdes B., Rejdali M., Achhal A. El Kadmiri, Jury J.L., Montserrat J.M. (eds.). Checklist of Vascular Plants of Morocco with identification keys. Consejo Superior de Investigaciones Cientificas. Madrid. 2: 555-558.
- Tjoelker M., Boratyński A., Bugała W. (red.). 2007. Biology and Ecology of Norway Spruce. Forestry Sciences 78. Springer, Dordrecht, the Netherandes.